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Section B. General Characters and Character States:
VIII. Growth and Development
[A. Growth Regions] [B. Growth Periodicity] [C. General Development] [D. Developmental Shoot Types]
A. Growth Regions
(Location of meristematic region on or within an organ or its parts)
- Adaxial or Abaxial. Growth region localized on top or bottom of a leaf part or any dorsi-ventral structure.
- Apical or Terminal. Growth region at the apex of the structure.
- Basal. Growth region at the base of a blade. as in grasses.
- Intercalary. Growth region near the base of an internode or base of blade.
- Interstitial. Growth all-over in an organ, no localized meristems, as in some fruits.
- Marginal. Growth region near the edge of a leaf.
B. Growth Periodicity
(Location of meristematic region on or within an organ or its parts)
- Accrescent. Growing after flowering or bud development has occurred, as the sepals in Hypericum and bud scales in Carya.
- Determinate. Growth of plant parts, the size of which is limited by cessation of meristematic activity during the year.
- Indeterminate or Evergrowing. Continual growth of plant parts, not limited by a cessation of meristematic activity.
- Intermittent. A renewal and cessation of meristematic activity which produces clusters of stems and/or leaves along an axis.
C. General Development
(Direction of maturation within the plant of its organs or its parts; e.g., from the base upward or
from the inside outward)
- Acropetal. Developing upward, toward apex.
- Basipetal. Developing downward, toward base.
- Centripetal. Developing from the outside inward, or from bottom upward.
- Centrifugal. Developing from the inside outward, or from top downward.
- Determinate. Central flower develops first, arresting development of primary axis.
- Indeterminate. Lateral flowers develop first with primary axis continuing to elongate or develop.
D. Developmental Shoot Types
(Classification based on origin and development of shoots)
- Dwarf Shoots or Spurs. Shoots that develop from preformed buds which have very short internodal lengths or intervals.
- Epicormic Shoots or Water Sprouts. Shoots that develop from dormant lateral buds on the trunk which have very long and frequently variable internodal lengths or intervals.
- Flushing Shoots. Shoots that develop from mature terminal buds several times during a season. Terminal bud will develop shoot with new terminal bud which will develop more shoots and a
terminal bud which will develop etc.--several times in a season with several flushes of growth.
- Heterophyllous Shoots. Shoots that develop from winter buds which do not contain the primordia of all the leaves to develop during the year.
- Lammas Shoots. Abnormal late season shoots that develop from the terminal bud, not a recurring phenomenon as in flushing shoots.
- Long Bud Shoots. Abnormal buds or shoots which elongate, then have arrested growth without the development of leaves and lateral branches.
- Long Shoots. Normal shoots that develop from terminal or axillary buds which have normal internodal lengths or intervals.
- Preformed Shoots. Normal shoots that develop from winter buds which contain primordia of all leaves that will expand during the season.
- Proleptic Shoots. Abnormal late season shoots that develop from the lateral buds immediately beneath the terminal.
- Sucker Shoots. Shoots that develop from adventitious buds on old stumps or roots, usually after cutting or injury, which have elongate internodal lengths and intervals.
- Syleptic Shoots. Abnormal shoots that develop from lateral buds before they have reached maturity.
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